$6.99 Buy It Now or Best Offer
free,30-Day Returns

Seller Store callistodesigns
(42696) 99.6%,

Location: Tucson, Arizona
Ships to: US,
Item: 361425539072

Restocking Fee:No
Return shipping will be paid by:Seller
All returns accepted:Returns Accepted
Item must be returned within:30 Days
Refund will be given as:Money Back

This listing is for a really fascinating trilobite specimen in a display jar, including an info card providing information about trilobites and the location this specimen came from. This kit is great for avid fossil and mineral specimen collectors or beginners. It would be a perfect gift set for getting someone interested in fossil collecting and science. The 1 centimeter scale cube is for size comparison only. It is not included in the sale. The photos are of several different specimens, but this listings is for one specimen with an info card. The photos show multiple specimens to give a representation of the variety of shapes and colors in these specimens. I offer a shipping discount for customers who combine their payments for multiple purchases into one payment! The discount is regular shipping price for the first item and just 50 cents for each additional item! To be sure you get your shipping discount just make sure all the items you want to purchase are in your cart. Auctions you win are added to your cart automatically. For any “buy it now” items or second chance offers, be sure to click the “add to cart” button, NOT the “buy it now” button. Once all of your items are in your cart just pay for them from your cart and the combined shipping discount should be applied automatically. I offer a money back guarantee on every item I sell. If you are not 100% happy with your purchase just send me a message to let me know and I will buy back the item for your full purchase price. Trilobites are extinct marine arthropods. They existed for 270 million years, from 520 to 250 million years ago. This makes them one of the most successful of all early animals, with 17,000 known species. The trilobites survived several extinction events but finally disappeared from the fossil record shortly before the Great Permian Extinction; Earth’s most severe mass extinction event, where 96% of all marine species went extinct. This specimen is Itagnostus interstrictus. It inhabited the ocean floor in what is now Utah, about 500 million years ago. If you have any questions, do not hesitate to ask me. TrilobiteFrom Wikipedia, the free encyclopediaJump to navigationJump to searchFor other uses, see Trilobite (disambiguation).TrilobitesTemporal range: 521–251.9 Ma PreЄЄOSDCPTJKPgNEarly Cambrian[1][2] – Late PermianKainops invius lateral and ventral.JPGKainops invius, early DevonianParadoxides sp.jpgParadoxides sp., late CambrianScientific classification editKingdom:AnimaliaPhylum:ArthropodaSubphylum:†TrilobitomorphaClass:†TrilobitaWalch, 1771[3]OrdersAgnostidaAsaphidaCorynexochidaHarpetidaNectaspidaRedlichiidaLichidaOdontopleuridaPhacopidaProetidaPtychopariidaTrilobites ( /ˈtraɪləˌbaɪt, ˈtrɪ-, -loʊ-/;[4][5] meaning “three lobes”) are a group of extinct marine arachnomorph arthropods that form the class Trilobita. Trilobites form one of the earliest-known groups of arthropods. The first appearance of trilobites in the fossil record defines the base of the Atdabanian stage of the Early Cambrian period (521 million years ago), and they flourished throughout the lower Paleozoic era before beginning a drawn-out decline to extinction when, during the Devonian, all trilobite orders except the Proetids died out. Trilobites disappeared in the mass extinction at the end of the Permian about 252 million years ago. The trilobites were among the most successful of all early animals, existing in oceans for over 270 million years.[6] By the time trilobites first appeared in the fossil record, they were already highly diversified and geographically dispersed. Because trilobites had wide diversity and an easily fossilized exoskeleton, they left an extensive fossil record, with some 17,000 known species spanning Paleozoic time. The study of these fossils has facilitated important contributions to biostratigraphy, paleontology, evolutionary biology, and plate tectonics. Trilobites are often placed within the arthropod subphylum Schizoramia within the superclass Arachnomorpha (equivalent to the Arachnata),[7] although several alternative taxonomies are found in the literature. Trilobites had many lifestyles; some moved over the sea bed as predators, scavengers, or filter feeders, and some swam, feeding on plankton. Most lifestyles expected of modern marine arthropods are seen in trilobites, with the possible exception of parasitism (where scientific debate continues).[8] Some trilobites (particularly the family Olenidae) are even thought to have evolved a symbiotic relationship with sulfur-eating bacteria from which they derived food.[9] Contents1Fossil record1.1Origins1.2Divergence and extinction1.2.1Evolutionary trends1.2.2Pre-Cambrian1.2.3Cambrian1.2.4Ordovician1.2.5Silurian and Devonian1.2.6Carboniferous and Permian1.2.7Final extinction2Fossil distribution3Importance4Taxonomy4.1Relationship to other taxa5Morphology5.1Terminology5.2Cephalon5.2.1Facial sutures5.2.1.1Dorsal sutures5.2.1.2Ventral sutures5.2.2Rostrum5.2.3Hypostome5.3Thorax5.4Pygidium5.5Prosopon (surface sculpture)5.6Spines6Soft body parts6.1Appendages6.2Digestive tract6.3Internal organs6.4Musculature7Sensory organs7.1Antennae7.2Eyes7.3Sensory pits8Development9History of usage and research10See also11References12Bibliography13External linksFossil record Redlichiida, such as this Paradoxides, may represent the ancestral trilobites Walliserops trifurcatus, from Jebel Oufatene mountain near Fezzou, MoroccoThe earliest trilobites known from the fossil record are redlichiids and ptychopariid bigotinids dated to some 540 to 520 million years ago.[1][2] Contenders for the earliest trilobites include Profallotaspis jakutensis (Siberia), Fritzaspis spp. (western USA), Hupetina antiqua (Morocco)[10] and Serrania gordaensis (Spain).[11] All trilobites are thought to have originated in present-day Siberia, with subsequent distribution and radiation from this location.[1] All Olenellina lack facial sutures (see below), and this is thought to represent the original state. The earliest sutured trilobite found so far (Lemdadella), occurs almost at the same time as the earliest Olenellina, suggesting the trilobites origin lies before the start of the Atdabanian, but without leaving fossils.[1] Other groups show secondary lost facial sutures, such as all Agnostina and some Phacopina. Another common feature of the Olenellina also suggests this suborder to be the ancestral trilobite stock: early protaspid stages have not been found, supposedly because these were not calcified, and this also is supposed to represent the original state.[12] Earlier trilobites may be found and could shed more light on the origin of trilobites.[1][13][14] Three specimens of a trilobite from Morocco, Megistaspis hammondi, dated 478 million years old contain fossilized soft parts.[15][16] OriginsEarly trilobites show all the features of the trilobite group as a whole; transitional or ancestral forms showing or combining the features of trilobites with other groups (e.g. early arthropods) do not seem to exist.[17] Morphological similarities between trilobites and early arthropod-like creatures such as Spriggina, Parvancorina, and other “trilobitomorphs” of the Ediacaran period of the Precambrian are ambiguous enough to make a detailed analysis of their ancestry complex.[12][18] Morphological similarities between early trilobites and other Cambrian arthropods (e.g. the Burgess shale fauna and the Maotianshan shales fauna) make analysis of ancestral relationships difficult as well.[19] Trilobites share a common ancestor with other arthropods before the Ediacaran-Cambrian boundary is still reasonable to assume. Evidence suggests that significant diversification had already occurred before trilobites were preserved in the fossil record, allowing for the “sudden” appearance of diverse trilobite groups with complex derived characteristics (e.g. eyes).[1][20] Divergence and extinction Olenoides serratus from the Mt. Stephen Trilobite Beds (Middle Cambrian) near Field, British Columbia, CanadaFor such a long-lasting group of animals, it is no surprise that trilobite evolutionary history is marked by a number of extinction events where some groups perished and surviving groups diversified to fill ecological niches with comparable or unique adaptations. Generally, trilobites maintained high diversity levels throughout the Cambrian and Ordovician periods before entering a drawn-out decline in the Devonian culminating in final extinction of the last few survivors at the end of the Permian period.[17] Evolutionary trendsPrincipal evolutionary trends from primitive morphologies, such as exemplified by Eoredlichia,[21] include the origin of new types of eyes, improvement of enrollment and articulation mechanisms, increased size of pygidium (micropygy to isopygy), and development of extreme spinosity in certain groups.[17] Changes also included narrowing of the thorax and increasing or decreasing numbers of thoracic segments.[21] Specific changes to the cephalon are also noted; variable glabella size and shape, position of eyes and facial sutures, and hypostome specialization.[21] Several morphologies appeared independently within different major taxa (e.g. eye reduction or miniaturization).[21] Effacement, the loss of surface detail in the cephalon, pygidium, or the thoracic furrows, is also a common evolutionary trend. Notable examples of this were the orders Agnostida and Asaphida, and the suborder Illaenina of the Corynexochida. Effacement is believed to be an indication of either a burrowing lifestyle or a pelagic one. Effacement poses a problem for taxonomists since the loss of details (particularly of the glabella) can make the determination of phylogenetic relationships difficult.[22] Pre-CambrianPhylogenetic biogeographic analysis of Early Cambrian Olenellidae and Redlichiidae suggests that a uniform trilobite fauna existed over Laurentia, Gondwana, and Siberia before the tectonic breakup of the supercontinent Pannotia between 600 and 550 million years ago.[1] Tectonic breakup of Pannotia then allowed for the diversification and radiation expressed later in the Cambrian as the distinctive olenellid province (Laurentia, Siberia, and Baltica) and the separate redlichid province (Australia, Antarctica, and China).[1][23] Breakup of Pannotia significantly antedates the first appearance of trilobites in the fossil record, supporting a long and cryptic development of trilobites extending perhaps as far back as 700 million years ago or possibly further back in time.[23] CambrianVery shortly after trilobite fossils appeared in the lower Cambrian, they rapidly diversified into the major orders that typified the Cambrian—Redlichiida, Ptychopariida, Agnostida, and Corynexochida. The first major crisis in the trilobite fossil record occurred in the Middle Cambrian; surviving orders developed isopygius or macropygius bodies and developed thicker cuticles, allowing better defense against predators (see Thorax below).[24] The end Cambrian mass extinction event marked a major change in trilobite fauna; almost all Redlichiida (including the Olenelloidea) and most Late Cambrian stocks became extinct.[17] A continuing decrease in Laurentian continental shelf area[25] is recorded at the same time as the extinctions, suggesting major environmental upheaval. Notable trilobite genera appearing in the Cambrian include:[26] Abadiella (Lower Cambrian)Judomia (Lower Cambrian)Olenellus (Lower Cambrian)Ellipsocephalus (Middle Cambrian)Elrathia (Middle Cambrian)Paradoxides (Middle Cambrian)Peronopsis (Middle Cambrian)Xiuqiella (Middle Cambrian)Yiliangella (Middle Cambrian)Yiliangellina (Middle Cambrian)Olenus (Late Cambrian)Ordovician Cast of Isotelus rex, the largest-known trilobite from the middle to upper Ordovician of North America Cheirurus sp., middle Ordovician age, Volkhov River, RussiaThe Early Ordovician is marked by vigorous radiations of articulate brachiopods, bryozoans, bivalves, echinoderms, and graptolites, with many groups appearing in the fossil record for the first time.[17] Although intra-species trilobite diversity seems to have peaked during the Cambrian,[27] trilobites were still active participants in the Ordovician radiation event with a new fauna taking over from the old Cambrian one.[28] Phacopida and Trinucleioidea are characteristic forms, highly differentiated and diverse, most with uncertain ancestors.[17] The Phacopida and other “new” clades almost certainly had Cambrian forebears, but the fact that they have avoided detection is a strong indication that novel morphologies were developing very rapidly.[12] Changes within the trilobite fauna during the Ordovician foreshadowed the mass extinction at the end of the Ordovician allowing many families to continue into the Silurian with little disturbance.[28] Ordovician trilobites were successful at exploiting new environments, notably reefs. The Ordovician mass extinction did not leave the trilobites unscathed; some distinctive and previously successful forms such as the Trinucleioidea and Agnostida became extinct. The Ordovician marks the last great diversification period amongst the trilobites: very few entirely new patterns of organisation arose post-Ordovician. Later evolution in trilobites was largely a matter of variations upon the Ordovician themes. By the Ordovician mass extinction, vigorous trilobite radiation has stopped and gradual decline beckons.[17] Some of the genera of Trilobites appearing in the Ordovician include:[26] Cyclopyge (Early to Late Ordovician)Selenopeltis (Early to Late Ordovician)Parabolina (Early Ordovician)Cheirurus (Middle Ordovician)Eodalmanitina (Middle Ordovician)Trinucleus (Middle Ordovician)Triarthrus (Late Ordovician)Silurian and Devonian Number of families, Middle Devonian to Upper Permian. Proetida – brown, Phacopida – steel blue, Lichida – clear blue, Harpetida, Odontopleurida and Corynexochida – pink, olive and purpleMost Early Silurian families constitute a subgroup of the Late Ordovician fauna. Few, if any, of the dominant Early Ordovician fauna survived to the end of the Ordovician, yet 74% of the dominant Late Ordovician trilobite fauna survived the Ordovician. Late Ordovician survivors account for all post-Ordovician trilobite groups except the Harpetida.[28] Silurian and Devonian trilobite assemblages are superficially similar to Ordovician assemblages, dominated by Lichida and Phacopida (including the well-known Calymenina). A number of characteristic forms do not extend far into the Devonian and almost all the remainder were wiped out by a series of dramatic Middle and Late Devonian extinctions.[21] Three orders and all but five families were exterminated by the combination of sea level changes and a break in the redox equilibrium (a meteorite impact has also been suggested as a cause).[21] Only a single order, the Proetida, survived into the Carboniferous.[17] Genera of trilobites during the Silurian and Devonian periods include:[26] Dalmanites (Early to Late Silurian)Calymene (Silurian)Encrinurus (Silurian)Exallaspis (Middle to Late Silurian)Paralejurus (Early Devonian)Lioharpes (Middle Devonian)Phacops (Middle to Late Devonian)Carboniferous and PermianThe Proetida survived for millions of years, continued through the Carboniferous period and lasted until the end of the Permian (when the vast majority of species on Earth were wiped out).[17] It is unknown why order Proetida alone survived the Devonian. The Proetida maintained relatively diverse faunas in deep water and shallow water, shelf environments throughout the Carboniferous.[21] For many millions of years the Proetida existed untroubled in their ecological niche.[17] An analogy would be today’s crinoids, which mostly exist as deep water species; in the Paleozoic era, vast ‘forests’ of crinoids lived in shallow near-shore environments.[17] Some of the genera of trilobites during the Carboniferous and Permian periods include:[26] Archegonus (Early to Middle Carboniferous)Hesslerides (Middle Carboniferous)Endops (Middle Permian)Triproetus (Late Carboniferous to Early Permian)Ditomopyge (Late Carboniferous to Late Permian)Final extinctionExactly why the trilobites became extinct is not clear; with repeated extinction events (often followed by apparent recovery) throughout the trilobite fossil record, a combination of causes is likely. After the extinction event at the end of the Devonian period, what trilobite diversity remained was bottlenecked into the order Proetida. Decreasing diversity[29] of genera limited to shallow-water shelf habitats, coupled with a drastic lowering of sea level (regression) meant that the final decline of trilobites happened shortly before the end of the Permian mass extinction event.[21] With so many marine species involved in the Permian extinction, the end of nearly 300 million successful years for the trilobite would not have been unexpected at the time.[29] Trilobites have no known direct descendants. Their closest living relatives would be the chelicerates. Though horseshoe crabs are often cited as their closest living relatives, they are no closer evolutionarily than other cheliceratans.[30] Fossil distribution Cruziana, fossil trilobite-burrowing trace A trilobite fragment (T) in a thin-section of an Ordovician limestone; E=echinoderm; scale bar is 2 mm Rusophycus, a “resting trace” of a trilobite; Ordovician of southern Ohio. Scale bar is 10 mm. Plate from Barrande’s work Système silurien du centre de la BohêmeTrilobites appear to have been exclusively marine organisms, since the fossilized remains of trilobites are always found in rocks containing fossils of other salt-water animals such as brachiopods, crinoids, and corals. Within the marine paleoenvironment, trilobites were found in a broad range from extremely shallow water to very deep water. Trilobites, like brachiopods, crinoids, and corals, are found on all modern continents, and occupied every ancient ocean from which Paleozoic fossils have been collected.[31] The remnants of trilobites can range from the preserved body to pieces of the exoskeleton, which it sheds in the process known as ecdysis. In addition, the tracks left behind by trilobites living on the sea floor are often preserved as trace fossils. There are three main forms of trace fossils associated with trilobites: Rusophycus; Cruziana & Diplichnites—such trace fossils represent the preserved life activity of trilobites active upon the sea floor. Rusophycus, the resting trace, are trilobite excavations involving little or no forward movement and ethological interpretations suggest resting, protection and hunting.[32] Cruziana, the feeding trace, are furrows through the sediment, which are believed to represent the movement of trilobites while deposit feeding.[33] Many of the Diplichnites fossils are believed to be traces made by trilobites walking on the sediment surface.[33] Care must be taken as similar trace fossils are recorded in freshwater[34] and post Paleozoic deposits,[35] representing non-trilobite origins. Trilobite fossils are found worldwide, with many thousands of known species. Because they appeared quickly in geological time, and moulted like other arthropods, trilobites serve as excellent index fossils, enabling geologists to date the age of the rocks in which they are found. They were among the first fossils to attract widespread attention, and new species are being discovered every year. In the United States, the best open to the public collection of trilobites is located in Hamburg, New York. Informally known as Penn Dixie, it was discovered in the 1970s by Dan Cooper. The shale quarry stopped mining in the 1960s,[36] but the amount of rock turnover showed large deposits of trilobites. As a well-known rock collector, he incited scientific and public interest in the location.[37] The fossils are dated to 350 million years ago when the Western New York Region was 30 degrees south of the equator and completely covered in water.[38] The site was purchased from Vincent Bonerb, Cavalcoli by the Town of Hamburg with the cooperation of the Hamburg Natural History Society to protect the land from development.[36] In 1994, the quarry became Penn Dixie Fossil Park & Nature Reserve when they received 501(c)3 status and was opened for visitation and collection of trilobite samples. The two most common found samples are Phacops rana and Greenops.[39] A famous location for trilobite fossils in the United Kingdom is Wren’s Nest, Dudley in the West Midlands, where Calymene blumenbachii is found in the Silurian Wenlock Group. This trilobite is featured on the town’s coat of arms and was named the Dudley Bug or Dudley Locust by quarrymen who once worked the now abandoned limestone quarries. Llandrindod Wells, Powys, Wales, is another famous trilobite location. The well-known Elrathia kingi trilobite is found in abundance in the Cambrian age Wheeler Shale of Utah.[40] Spectacularly preserved trilobite fossils, often showing soft body parts (legs, gills, antennae, etc.) have been found in British Columbia, Canada (the Cambrian Burgess Shale and similar localities); New York, U.S.A. (Ordovician Walcott–Rust quarry, near Russia, and Beecher’s Trilobite Bed, near Rome); China (Lower Cambrian Maotianshan Shales near Chengjiang); Germany (the Devonian Hunsrück Slates near Bundenbach) and, much more rarely, in trilobite-bearing strata in Utah (Wheeler Shale and other formations), Ontario, and Manuels River, Newfoundland and Labrador. Sites in Morocco also yield very well preserved trilobites. The French palaeontologist Joachim Barrande (1799–1883) carried out his landmark study of trilobites in the Cambrian, Ordovician and Silurian of Bohemia, publishing the first volume of Système silurien du centre de la Bohême in 1852. ImportanceThe study of Paleozoic trilobites in the Welsh-English borders by Niles Eldredge was fundamental in formulating and testing punctuated equilibrium as a mechanism of evolution.[41][42][43] Identification of the ‘Atlantic’ and ‘Pacific’ trilobite faunas in North America and Europe[44] implied the closure of the Iapetus Ocean (producing the Iapetus suture),[45] thus providing important supporting evidence for the theory of continental drift.[46][47] Trilobites have been important in estimating the rate of speciation during the period known as the Cambrian explosion because they are the most diverse group of metazoans known from the fossil record of the early Cambrian.[48][49] Trilobites are excellent stratigraphic markers of the Cambrian period: researchers who find trilobites with alimentary prosopon, and a micropygium, have found Early Cambrian strata.[50] Most of the Cambrian stratigraphy is based on the use of trilobite marker fossils.[51][52][53] Trilobites are the state fossils of Ohio (Isotelus), Wisconsin (Calymene celebra) and Pennsylvania (Phacops rana). TaxonomySee also: List of trilobite generaThe taxonomic rank of the many groupings of trilobites can be derived from the suffix to its name. The following table lists the different ranks (from widest to closest relatives), with the corresponding suffix and an example. suffixes to the names of taxonomic trilobite groupings of different rankranksuffixexampleclass-aTrilobitaorder-idaPtychopariidasuborder-inaEodiscinasuperfamily-oideaAcastoideafamily-idaeCyclopygidaesubfamily-inaeCybelinaeThe 10 trilobite orders are Agnostida, Redlichiida, Corynexochida, Lichida, Odontopleurida, Phacopida, Proetida, Asaphida, Harpetida and Ptychopariida. Sometimes the Nektaspida are included, but these lack a calcified exoskeleton and eyes. Other scholars do not consider Agnostina to be trilobites, and hence not related to the Eodiscina. This latter suborder should than be elevated and be called Eodiscida. Despite their rich fossil record with thousands of described genera found throughout the world, the taxonomy and phylogeny of trilobites have many uncertainties.[54] Except possibly for the members of the orders Phacopida and Lichida (which first appear during the early Ordovician), nine of the eleven trilobite orders appear prior to the end of the Cambrian. Most scientists believe that order Redlichiida, more specifically its suborder Redlichiina, contains a common ancestor of all other orders, with the possible exception of the Agnostina. While many potential phylogenies are found in the literature, most have suborder Redlichiina giving rise to orders Corynexochida and Ptychopariida during the Lower Cambrian, and the Lichida descending from either the Redlichiida or Corynexochida in the Middle Cambrian. Order Ptychopariida is the most problematic order for trilobite classification. In the 1959 Treatise on Invertebrate Paleontology,[55] what are now members of orders Ptychopariida, Asaphida, Proetida and Harpetida were grouped together as order Ptychopariida; subclass Librostoma was erected in 1990[56] to encompass all of these orders, based on their shared ancestral character of a natant (unattached) hypostome. The most recently recognized of the nine trilobite orders, Harpetida, was erected in 2002.[57] The progenitor of order Phacopida is unclear. Relationship to other taxaOnce soft-part anatomy had been recovered, the trilobites were originally allied to the Crustacea. A relationship with the Chelicerata, in a clade termed Arachnomorpha (Arachnata), was in vogue for some time,[58]but, a more recent analysis of Panarthropoda suggests the trilobites are one of two major branches of Artiopoda.[30] MorphologyWhen trilobites are found, only the exoskeleton is preserved (often in an incomplete state) in all but a handful of locations. A few locations (Lagerstätten) preserve identifiable soft body parts (legs, gills, musculature & digestive tract) and enigmatic traces of other structures (e.g. fine details of eye structure) as well as the exoskeleton. Trilobites range in length from minute (less than 3 millimetres (0.12 in)) to very large (over 30 centimetres (12 in)), with an average size range of 3–10 cm (1.2–3.9 in). Supposedly the smallest species is Acanthopleurella stipulae with a maximum of 1.5 millimetres (0.059 in).[59] The world’s largest-known trilobite specimen, assigned to Isotelus rex of 72 cm, was found in 1998 by Canadian scientists in Ordovician rocks on the shores of Hudson Bay.[25] The trilobite body is divided into three major sections (tagmata): 1 – cephalon; 2 – thorax; 3 – pygidium. Trilobites are so named for the three longitudinal lobes: 4 – right pleural lobe; 5 – axial lobe; 6 – left pleural lobe; the antennae and legs are not shown in these diagramsThe exoskeleton is composed of calcite and calcium phosphate minerals in a lattice of chitin that covers the upper surface (dorsal) of the trilobite and curled round the lower edge to produce a small fringe called the “doublure”. Three distinctive tagmata (sections) are present: cephalon (head); thorax (body) and pygidium (tail). TerminologyAs might be expected for a group of animals comprising c. 5,000 genera,[60] the morphology and description of trilobites can be complex. Despite morphological complexity and an unclear position within higher classifications, there are a number of characteristics which distinguish the trilobites from other arthropods: a generally sub-elliptical, dorsal, chitinous exoskeleton divided longitudinally into three distinct lobes (from which the group gets its name); having a distinct, relatively large head shield (cephalon) articulating axially with a thorax comprising articulated transverse segments, the hindmost of which are almost invariably fused to form a tail shield (pygidium). When describing differences between trilobite taxa, the presence, size, and shape of the cephalic features are often mentioned. During moulting, the exoskeleton generally splits between the head and thorax, which is why so many trilobite fossils are missing one or the other. In most groups facial sutures on the cephalon helped facilitate moulting. Similar to lobsters and crabs, trilobites would have physically “grown” between the moult stage and the hardening of the new exoskeleton. CephalonSee also: Cephalon (arthropod anatomy)Morphology of the trilobite cephalonLabeled diagram of major parts of the cephalonThe major subdivisions of the cephalon The subdivisions can be further broken down into different areas used in describing trilobite cephalic morphology. 1 – preocular area; 2 – palpebral area; 3 – postocular area; 4 – posterolateral projection; 5 – occipital ring; 6 – glabella; 7 – posterior area; 8 – lateral border; 9 – librigenal area; 10 – preglabellar areaA trilobite’s cephalon, or head section, is highly variable with a lot of morphological complexity. The glabella forms a dome underneath which sat the “crop” or “stomach”. Generally the exoskeleton has few distinguishing ventral features, but the cephalon often preserves muscle attachment scars and occasionally the hypostome, a small rigid plate comparable to the ventral plate in other arthropods. A toothless mouth and stomach sat upon the hypostome with the mouth facing backward at the rear edge of the hypostome. Hypostome morphology is highly variable; sometimes supported by an un-mineralised membrane (natant), sometimes fused onto the anterior doublure with an outline very similar to the glabella above (conterminant) or fused to the anterior doublure with an outline significantly different from the glabella (impendent). Many variations in shape and placement of the hypostome have been described.[56] The size of the glabella and the lateral fringe of the cephalon, together with hypostome variation, have been linked to different lifestyles, diets and specific ecological niches.[61] The anterior and lateral fringe of the cephalon is greatly enlarged in the Harpetida, in other species a bulge in the pre-glabellar area is preserved that suggests a brood pouch.[62] Highly complex compound eyes are another obvious feature of the cephalon. Facial suturesFacial or cephalic sutures are the natural fracture lines in the cephalon of trilobites. Their function was to assist the trilobite in shedding its old exoskeleton during ecdysis (or molting).[63] All species assigned to the suborder Olenellina, that became extinct at the very end of the Early Cambrian (like Fallotaspis, Nevadia, Judomia, and Olenellus) lacked facial sutures. They are believed to have never developed facial sutures, having pre-dated their evolution. Because of this (along with other primitive characteristics), they are thought to be the earliest ancestors of later trilobites.[64][1] Some other later trilobites also lost facial sutures secondarily.[64] The type of sutures found in different species are used extensively in the taxonomy and phylogeny of trilobites.[65] Dorsal suturesThe dorsal surface of the trilobite cephalon (the frontmost tagma, or the ‘head’) can be divided into two regions—the cranidium and the librigena (“free cheeks”). The cranidium can be further divided into the glabella (the central lobe in the cephalon) and the fixigena (“fixed cheeks”).[66] The facial sutures lie along the anterior edge, at the division between the cranidium and the librigena. Trilobite facial sutures on the dorsal side can be roughly divided into five main types according to where the sutures end relative to the genal angle (the edges where the side and rear margins of the cephalon converge).[67] Absent – Facial sutures are lacking in the Olenellina. This is considered a primitive state, and is always combined with the presence of eyes.Proparian – The facial suture ends in front of the genal angle, along the lateral margin.[66] Example genera showing this type of suture include Dalmanites of Phacopina (Phacopida) and Ekwipagetia of Eodiscina (Agnostida).Gonatoparian – The facial suture ends at the tip of the genal angle.[68] Example genera showing this type of suture include Calymene and Trimerus of Calymenina (Phacopida).[65]Opisthoparian – The facial suture ends at the posterior margin of the cephalon.[65] Example genera showing this type of suture include Peltura of Olenina (Ptychopariida) and Bumastus of Illaenina (Corynexochida). This is the most common type of facial suture.[65]Hypoparian or marginal – In some trilobites, dorsal sutures may be secondarily lost. Several exemplary time series of species show the “migration” of the dorsal suture until it coincides with the margins of the cephalon.[66] As the visual surface of the eye is on the diminishing free cheek (or librigena), the number of lenses tends to go down, and eventually the eye disappears. The loss of dorsal sutures may arise from the proparian state, such as in some Eodiscina like Weymouthia, all Agnostina, and some Phacopina such as Ductina. The marginal sutures exhibited by the harpetids and trinucleioids, are derived from opisthoparian sutures.[69] On the other hand, blindness is not always accompanied by the loss of facial sutures.Trilobite facial suture types.pngThe primitive state of the dorsal sutures is proparian. Opisthoparian sutures have developed several times independently. There are no examples of proparian sutures developing in taxa with opisthoparian ancestry. Trilobites that exhibit opisthoparian sutures as adults commonly have proparian sutures as instars (known exceptions being Yunnanocephalus and Duyunaspis).[70] Hypoparian sutures have also arisen independently in several groups of trilobites. The course of the facial sutures from the front of the visual surface varies at least as strongly as it does in the rear, but the lack of a clear reference point similar to the genal angle makes it difficult to categorize. One of the more pronounced states is that the front of the facial sutures do not cut the lateral or frontal border on its own, but coincide in front of the glabella, and cut the frontal border at the midline. This is, inter alia, the case in the Asaphida. Even more pronounced is the situation that the frontal branches of the facial sutures end in each other, resulting in yoked free cheeks. This is known in Triarthrus, and in the Phacopidae, but in that family are the facial sutures not functional, as can be concluded from the fact that free cheeks are not found separated from the cranidium. There are also two types of sutures in the dorsal surface connected to the compound eyes of trilobites.[65][71] They are: Ocular sutures – are sutures surrounding the edges of the compound eye. Trilobites with these sutures lose the entire surface of the eyes when molting. It is common among Cambrian trilobites.Palpebral sutures – are sutures which form part of the dorsal facial suture running along the top edges of the compound eye.Ventral suturesTrilobite cephalon ventral anatomy.pngDorsal facial sutures continue downward to the ventral side of the cephalon where they become the Connective sutures that divide the doublure. The following are the types of ventral sutures.[71] Connective sutures – are the sutures that continue from the facial sutures past the front margin of the cephalon.Rostral suture – is only present when the trilobite possesses a rostrum (or rostral plate). It connects the rostrum to the front part of the dorsal cranidium.Hypostomal suture – separates the hypostome from the doublure when the hypostome is of the attached type. It is absent when the hypostome is free-floating (i.e. natant). it is also absent in some coterminant hypostomes where the hypostome is fused to the doublure.Median suture – exhibited by asaphid trilobites, they are formed when instead of becoming connective sutures, the two dorsal sutures converge at a point in front of the cephalon then divide straight down the center of the doublure.RostrumThe rostrum (or the rostral plate) is a distinct part of the doublure located at the front of the cephalon. It is separated from the rest of the doublure by the rostral suture. During molting in trilobites like Paradoxides, the rostrum is used to anchor the front part of the trilobite as the cranidium separates from the librigena. The opening created by the arching of the body provides an exit for the molting trilobite. It is absent in some trilobites like Lachnostoma. HypostomeFurther information: Hypostome (trilobite) Asaphus expansus ventral side prepared, showing the attachment of the hypostomeThe hypostome is the hard mouthpart of the trilobite found on the ventral side of the cephalon typically below the glabella. The hypostome can be classified into three types based on whether they are permanently attached to the rostrum or not and whether they are aligned to the anterior dorsal tip of the glabella. Natant – Hypostome not attached to doublure. Aligned with front edge of glabella.Conterminant – Hypostome attached to rostral plate of doublure. Aligned with front edge of glabella.Impendent – Hypostome attached to rostral plate but not aligned to glabella.Below is an illustration of the three types. The doublure is shown in light gray, the inside surface of the cephalon in dark gray, and the hypostome in light blue. The glabella is outlined in red broken lines. Trilobite hypostome types based on attachment (labeled).pngThorax An enrolled phacopid trilobite Flexiclaymene meeki from the Upper Ordovician of OhioThe thorax is a series of articulated segments that lie between the cephalon and pygidium. The number of segments varies between 2 and 103[72] with most species in the 2 to 16 range.[59] Each segment consists of the central axial ring and the outer pleurae, which protected the limbs and gills. The pleurae are sometimes abbreviated or extended to form long spines. Apodemes are bulbous projections on the ventral surface of the exoskeleton to which most leg muscles attached, although some leg muscles attached directly to the exoskeleton.[73] Determining a junction between thorax and pygidium can be difficult and many segment counts suffer from this problem.[59] Trilobite fossils are often found “enrolled” (curled up) like modern pill bugs for protection; evidence suggests enrollment helped protect against the inherent weakness of the arthropod cuticle that was exploited by anomalocarid predators.[24] Some trilobites achieved a fully closed capsule (e.g. Phacops), while others with long pleural spines (e.g. Selenopeltis) left a gap at the sides or those with a small pygidium (e.g. Paradoxides) left a gap between the cephalon and pygidium.[59] In Phacops, the pleurae overlap a smooth bevel (facet) allowing a close seal with the doublure.[73] The doublure carries a Panderian notch or protuberance on each segment to prevent over rotation and achieve a good seal.[73] Even in an agnostid, with only 2 articulating thoracic segments, the process of enrollment required a complex musculature to contract the exoskeleton and return to the flat condition.[74] PygidiumThe pygidium is formed from a number of segments and the telson fused together. Segments in the pygidium are similar to the thoracic segments (bearing biramous limbs) but are not articulated. Trilobites can be described based on the pygidium being micropygous (pygidium smaller than cephalon), subisopygous (pygidium sub equal to cephalon), isopygous (pygidium equal in size to cephalon), or macropygous (pygidium larger than cephalon). Prosopon (surface sculpture) Koneprusia brutoni, an example of a species with elaborate spines from the Devonian Hamar Laghdad Formation, Alnif, MoroccoTrilobite exoskeletons show a variety of small-scale structures collectively called prosopon. Prosopon does not include large scale extensions of the cuticle (e.g. hollow pleural spines) but to finer scale features, such as ribbing, domes, pustules, pitting, ridging and perforations. The exact purpose of the prosopon is not resolved but suggestions include structural strengthening, sensory pits or hairs, preventing predator attacks and maintaining aeration while enrolled.[59] In one example, alimentary ridge networks (easily visible in Cambrian trilobites) might have been either digestive or respiratory tubes in the cephalon and other regions.[17] SpinesSome trilobites such as those of the order Lichida evolved elaborate spiny forms, from the Ordovician until the end of the Devonian period. Examples of these specimens have been found in the Hamar Laghdad Formation of Alnif in Morocco. There is a serious counterfeiting and fakery problem with much of the Moroccan material that is offered commercially. Spectacular spined trilobites have also been found in western Russia; Oklahoma, USA; and Ontario, Canada. Some trilobites had horns on their heads similar to several modern beetles. Based on the size, location, and shape of the horns it has been suggested that these horns may have been used to combat for mates. Horns were widespread in the Raphiophoridae family (Asaphida).[75] Another function of these spines was protection from predators. When enrolled, trilobite spines offered additional protection. This conclusion is likely to be applicable to other trilobites as well, such as in the Phacopid trilobite genus Walliserops, that developed spectacular tridents.[76] Soft body partsOnly 21 or so species are described from which soft body parts are preserved,[73][77] so some features (e.g. the posterior antenniform cerci preserved only in Olenoides serratus)[78] remain difficult to assess in the wider picture.[79] life reconstruction of Triarthrus eatoni based on preserved soft body partsAppendages Drawing of a biramous leg of Agnostus pisiformis Ventral side of Triarthrus eatoni, showing antennas and biramous legs preserved in pyrite An exceptionally well preserved Olenoides serratus from the Burgess Shale. The antennae and legs are preserved as reflective carbon film.Trilobites had a single pair of preoral antennae and otherwise undifferentiated biramous limbs (2, 3 or 4 cephalic pairs, followed by one pair per thoracic segment and some pygidium pairs).[73][77] Each endopodite (walking leg) had 6 or 7 segments,[77] homologous to other early arthropods.[79] Endopodites are attached to the coxa, which also bore a feather-like exopodite, or gill branch, which was used for respiration and, in some species, swimming.[79] The inside of the coxa (or gnathobase) carries spines, probably to process prey items.[80] The last exopodite segment usually had claws or spines.[73] Many examples of hairs on the legs suggest adaptations for feeding (as for the gnathobases) or sensory organs to help with walking.[79] Digestive tractThe toothless mouth of trilobites was situated on the rear edge of the hypostome (facing backward), in front of the legs attached to the cephalon. The mouth is linked by a small esophagus to the stomach that lay forward of the mouth, below the glabella. The “intestine” led backward from there to the pygidium.[73] The “feeding limbs” attached to the cephalon are thought to have fed food into the mouth, possibly “slicing” the food on the hypostome and/or gnathobases first. Alternative lifestyles are suggested, with the cephalic legs used to disturb the sediment to make food available. A large glabella, (implying a large stomach), coupled with an impendent hypostome has been used as evidence of more complex food sources, i.e. possibly a carnivorous lifestyle.[8] Internal organsWhile there is direct and implied evidence for the presence and location of the mouth, stomach and digestive tract (see above) the presence of heart, brain and liver are only implied (although “present” in many reconstructions) with little direct geological evidence.[79] MusculatureAlthough rarely preserved, long lateral muscles extended from the cephalon to midway down the pygidium, attaching to the axial rings allowing enrollment while separate muscles on the legs tucked them out of the way.[73] Sensory organsMany trilobites had complex eyes; they also had a pair of antennae. Some trilobites were blind, probably living too deep in the sea for light to reach them. As such, they became secondarily blind in this branch of trilobite evolution. Other trilobites (e.g., Phacops rana and Erbenochile erbeni) had large eyes that were for use in well lit, predator-filled waters. AntennaeThe pair of antennae suspected in most trilobites (and preserved in a few examples) were highly flexible to allow them to be retracted when the trilobite was enrolled. One species (Olenoides serratus) preserves antenna-like cerci, which project from the rear of the trilobite.[78] EyesEven the earliest trilobites had complex, compound eyes with lenses made of calcite (a characteristic of all trilobite eyes), confirming that the eyes of arthropods and probably other animals could have developed before the Cambrian.[20] Improving eyesight of both predator and prey in marine environments has been suggested as one of the evolutionary pressures furthering an apparent rapid development of new life forms during what is known as the Cambrian explosion.[81] Trilobite eyes were typically compound, with each lens being an elongated prism.[82] The number of lenses in such an eye varied: some trilobites had only one, while some had thousands of lenses in a single eye. In compound eyes, the lenses were typically arranged hexagonally.[17] The fossil record of trilobite eyes is complete enough that their evolution can be studied through time, which compensates to some extent the lack of preservation of soft internal parts.[83] Lenses of trilobites’ eyes were made of calcite (calcium carbonate, CaCO3). Pure forms of calcite are transparent, and some trilobites used crystallographically oriented, clear calcite crystals to form each lens of each eye.[84] Rigid calcite lenses would have been unable to accommodate to a change of focus like the soft lens in a human eye would; in some trilobites, the calcite formed an internal doublet structure,[85] giving superb depth of field and minimal spherical aberration, according to optical principles discovered by French scientist René Descartes and Dutch physicist Christiaan Huygens in the 17th century.[82][85] A living species with similar lenses is the brittle star Ophiocoma wendtii.[86] In other trilobites, with a Huygens interface apparently missing, a gradient-index lens is invoked with the refractive index of the lens changing toward the center.[87] Sublensar sensory structures have been found in the eyes of some phacopid trilobites.[88] The structures consist of what appear to be several sensory cells surrounding a rhadomeric structure, resembling closely the sublensar structures found in the eyes of many modern arthropod apposition eyes, especially Limulus, a genus of horseshoe crabs.[88] Holochroal eyes had a great number (sometimes over 15,000) of small (30–100 μm, rarely larger)[83] lenses. Lenses were hexagonally close packed, touching each other, with a single corneal membrane covering all lenses.[84] Each lens was in direct contact with adjacent lenses. Holochroal eyes are the ancestral eye of trilobites, and are by far the most common, found in all orders except the Agnostida, and through the entirety of the Trilobites’ existence.[83] Little is known of the early history of holochroal eyes; Lower and Middle Cambrian trilobites rarely preserve the visual surface.[83] The spatial resolving power of grated eyes (such as holochroal eyes) is dependent on light intensity, circular motion, receptor density, registered light angle, and the extent to which the signal of individual rhabdoms are neurally combined. This implies that lenses need to be larger under low light conditions (such as for Pricyclopyge, when comparing it to Carolinites), and for fast moving predators and prey. As the circular velocity caused by the forward speed of an animal itself is much higher for the ommatidia directed perpendicular to the movement, fast-moving trilobites (such as Carolinites) have eyes flattened from the side and more curved were ommatia are directed to the front or back. Thus eye morphology can be used to make assumptions about the ecosystem of trilobites.[89] The schizochroal eye of Erbenochile erbenii; the eye shade is unequivocal evidence that some trilobites were diurnal[90]Schizochroal eyes typically had fewer (around 700), larger lenses than holochroal eyes and are found only in Phacopina. Each lens had a cornea, and adjacent lenses were separated by thick interlensar cuticle, known as sclera. Schizochroal eyes appear quite suddenly in the early Ordovician, and were presumably derived from a holochroal ancestor.[83] Field of view (all-around vision), eye placement and coincidental development of more efficient enrollment mechanisms point to the eye as a more defensive “early warning” system than directly aiding in the hunt for food.[83] Modern eyes that are functionally equivalent to the schizochroal eye were not thought to exist,[84] but are found in the modern insect species Xenos peckii.[91]Abathochroal eyes are found only in Cambrian Eodiscina, and have around 70 small separate lenses that had individual cornea.[92] The sclera was separate from the cornea, and was not as thick as the sclera in schizochroal eyes.[84] Although well preserved examples are sparse in the early fossil record, abathochroal eyes have been recorded in the lower Cambrian, making them among the oldest known.[84] Environmental conditions seem to have resulted in the later loss of visual organs in many Eodiscina.[84]Secondary blindness is not uncommon, particularly in long lived groups such as the Agnostida and Trinucleioidea. In Proetida and Phacopina from western Europe and particularly Tropidocoryphinae from France (where there is good stratigraphic control), there are well studied trends showing progressive eye reduction between closely related species that eventually leads to blindness.[84] Several other structures on trilobites have been explained as photo-receptors.[84] Of particular interest are “macula”, the small areas of thinned cuticle on the underside of the hypostome. In some trilobites macula are suggested to function as simple “ventral eyes” that could have detected night and day or allowed a trilobite to navigate while swimming (or turned) upside down.[87] Pitted fringes on the cephalon of Trinucleoidea trilobiteSensory pitsThere are several types of prosopon that have been suggested as sensory apparatus collecting chemical or vibrational signals. The connection between large pitted fringes on the cephalon of Harpetida and Trinucleoidea with corresponding small or absent eyes makes for an interesting possibility of the fringe as a “compound ear”.[84] Development Five Stages in the development of Sao hirsutaTrilobites grew through successive moult stages called instars, in which existing segments increased in size and new trunk segments appeared at a sub-terminal generative zone during the anamorphic phase of development. This was followed by the epimorphic developmental phase, in which the animal continued to grow and moult, but no new trunk segments were expressed in the exoskeleton. The combination of anamorphic and epimorphic growth constitutes the hemianamorphic developmental mode that is common among many living arthropods.[93] Trilobite development was unusual in the way in which articulations developed between segments, and changes in the development of articulation gave rise to the conventionally recognized developmental phases of the trilobite life cycle (divided into 3 stages), which are not readily compared with those of other arthropods. Actual growth and change in external form of the trilobite would have occurred when the trilobite was soft shelled, following moulting and before the next exoskeleton hardened.[94] Elrathia kingii growth series with holaspids ranging from 16.2 mm to 39.8 mm in lengthTrilobite larvae are known from the Cambrian to the Carboniferous[95] and from all sub-orders.[94][96] As instars from closely related taxa are more similar than instars from distantly related taxa, trilobite larvae provide morphological information important in evaluating high-level phylogenetic relationships among trilobites.[94] Despite the absence of supporting fossil evidence, their similarity to living arthropods has led to the belief that trilobites multiplied sexually and produced eggs.[94][97] Some species may have kept eggs or larvae in a brood pouch forward of the glabella,[62] particularly when the ecological niche was challenging to larvae.[9] Size and morphology of the first calcified stage are highly variable between (but not within) trilobite taxa, suggesting some trilobites passed through more growth within the egg than others. Early developmental stages prior to calcification of the exoskeleton are a possibility (suggested for fallotaspids),[12] but so is calcification and hatching coinciding.[94] The earliest post-embryonic trilobite growth stage known with certainty are the “protaspid” stages (anamorphic phase).[94] Starting with an indistinguishable proto-cephalon and proto-pygidium (anaprotaspid) a number of changes occur ending with a transverse furrow separating the proto-cephalon and proto-pygidium (metaprotaspid) that can continue to add segments. Segments are added at the posterior part of the pygidium, but all segments remain fused together.[94][96] The “meraspid” stages (anamorphic phase) are marked by the appearance of an articulation between the head and the fused trunk. Prior to the onset of the first meraspid stage the animal had a two-part structure—the head and the plate of fused trunk segments, the pygidium. During the meraspid stages, new segments appeared near the rear of the pygidium as well as additional articulations developing at the front of the pygidium, releasing freely articulating segments into the thorax. Segments are generally added one per moult (although two per moult and one every alternate moult are also recorded), with number of stages equal to the number of thoracic segments. A substantial amount of growth, from less than 25% up to 30%–40%, probably took place in the meraspid stages.[94] The “holaspid” stages (epimorphic phase) commence when a stable, mature number of segments has been released into the thorax. Moulting continued during the holaspid stages, with no changes in thoracic segment number.[94] Some trilobites are suggested to have continued moulting and growing throughout the life of the individual, albeit at a slower rate on reaching maturity. Some trilobites showed a marked transition in morphology at one particular instar, which has been called “trilobite metamorphosis”. Radical change in morphology is linked to the loss or gain of distinctive features that mark a change in mode of life.[98] A change in lifestyle during development has significance in terms of evolutionary pressure, as the trilobite could pass through several ecological niches on the way to adult development and changes would strongly affect survivorship and dispersal of trilobite taxa.[94] It is worth noting that trilobites with all protaspid stages solely planktonic and later meraspid stages benthic (e.g. asaphids) failed to last through the Ordovician extinctions, while trilobites that were planktonic for only the first protaspid stage before metamorphosing into benthic forms survived (e.g. lichids, phacopids).[98] Pelagic larval life-style proved ill-adapted to the rapid onset of global climatic cooling and loss of tropical shelf habitats during the Ordovician.[25] History of usage and research Drawing of Ogygiocarella debuchii by Rev. Edward Lhwyd, made in 1698Rev. Edward Lhwyd published in 1698 in The Philosophical Transactions of the Royal Society, the oldest scientific journal in the English language, part of his letter “Concerning Several Regularly Figured Stones Lately Found by Him”, that was accompanied by a page of etchings of fossils.[99] One of his etchings figured a trilobite he found near Llandeilo, probably on the grounds of Lord Dynefor’s castle, he described as “… the skeleton of some flat Fish …”.[2] The discovery of Calymene blumenbachii (the Dudley locust) in 1749 by Charles Lyttleton, could be identified as the beginning of trilobite research. Lyttleton submitted a letter to the Royal Society of London in 1750 concerning a “petrified insect” he found in the “limestone pits at Dudley”. In 1754, Manuel Mendez da Costa proclaimed that the Dudley locust was not an insect, but instead belonged to “the crustaceous tribe of animals.” He proposed to call the Dudley specimens Pediculus marinus major trilobos (large trilobed marine louse), a name which lasted well into the 1800s. German naturalist Johann Walch, who executed the first inclusive study of this group, proposed the use of the name “trilobite”. He considered it appropriate to derive the name from the unique three-lobed character of the central axis and a pleural zone to each side.[100] Written descriptions of trilobites date possibly from the third century BC and definitely from the fourth century AD. The Spanish geologists Eladio Liñán and Rodolfo Gozalo argue that some of the fossils described in Greek and Latin lapidaries as scorpion stone, beetle stone, and ant stone, refer to trilobite fossils. Less ambiguous references to trilobite fossils can be found in Chinese sources. Fossils from the Kushan formation of northeastern China were prized as inkstones and decorative pieces.[99] In the New World, American fossil hunters found plentiful deposits of Elrathia kingi in western Utah in the 1860s. Until the early 1900s, the Ute Native Americans of Utah wore these trilobites, which they called pachavee (little water bug), as amulets. A hole was bored in the head and the fossil was worn on a string.[101] According to the Ute themselves, trilobite necklaces protect against disease and bullets. In 1931, Frank Beckwith uncovered evidence of the Ute use of trilobites. Travelling through the badlands, he photographed two petroglyphs that most likely represent trilobites. On the same trip he examined a burial, of unknown age, with a drilled trilobite fossil lying in the chest cavity of the interred. Since then, trilobite amulets have been found all over the Great Basin, as well as in British Columbia and Australia.[99] In the 1880s, archaeologists discovered in the Grotte du Trilobite (Caves of Arcy-sur-Cure, Yonne, France) a much-handled trilobite fossil that had been drilled as if to be worn as a pendant. The occupation stratum in which the trilobite was found has been dated as fifteen thousand years old. Because the pendant was handled so much, the species of trilobite cannot be determined. This type of trilobite is not found around Yonne, so it may have been highly prized and traded from elsewhere.[99] See alsoiconPaleontology portaliconArthropods portalNotostracaHorseshoe crabIsopodaList of trilobite generaEvolutionary history of life

Frequently Asked Questions About Trilobite ITAGNOSTUS Specimen Fossil Mineral Sea Life WHEELR SHALE w/ ID Card in My Website

swidnw.com is the best online shopping platform where you can buy Trilobite ITAGNOSTUS Specimen Fossil Mineral Sea Life WHEELR SHALE w/ ID Card from renowned brand(s). swidnw.com delivers the most unique and largest selection of products from across the world especially from the US, UK and India at best prices and the fastest delivery time.

What are the best-selling Trilobite ITAGNOSTUS Specimen Fossil Mineral Sea Life WHEELR SHALE w/ ID Card on swidnw.com?

swidnw.com helps you to shop online and delivers Moschino to your doorstep. The best-selling Moschino on swidnw.com are: Moschino MOS020/S Womens Large Metal Sunglass DDB3X Gold Copper w Pink 59-16 Love Moschino Purse Borsa Vit Black Genuine Leather Gold Studded Shoulder Bag Moschino Brown Faux Pearl Eyeglasses *FRAMES ONLY* MO084-04 53-16-135 Italy NWOT Moschino Classic Body Powder With Puff New Moschino Women’s 8 Off White Linen Blend Relaxed Pants MOSCHINO COUTURE! Black Classic Shift Dress – Gold Grommets USA 8 GB 10 Moschino by Moschino, 2.5 oz EDT Spray for Women Women’s Moschino Black Tapered Leg Jersey Knit Pants Size 10 Moschino White X Kelloggs Tony The Tiger Graphic Sweatshirt LOVE MOSCHINO Womens Dress Size 46 Size 10 Python Heart Print Logo Long Sleeve Moschino Cheap And Chic Tweed Style 100% Wool Blazer Jacket UK Size 10 Moschino Cheap & Chic Dress Mixed Print Silk Twill Size US 4 Twist-Front Sheath Moschino Cheap & Chic Black Jacket Size 46 IT Moschino Boutique Tweed Jacket Pink Cropped Pearl Buttons Long Sleeves Size 10 M VINTAGE MOSCHINO RARE ESTATE SALE FIND BLACK CAMEO PATCH PULL ON HI LO SKIRT S/M AW20 Moschino Couture Jeremy Scott Shipping Boxes Packaging Materials Overcoat MOSCHINO LEATHER SKIRT Moschino Jeans Women’s Linen Blazer Size 6 Navy Blue Dark Academia Boutique Moschino stretch High Gather Waist skirt Size IT40 USA6 F36 8 Moschino Cheap Chic Eyelet Wiggle Dress White Cotton Italy Boutique Moschino! -Print Ribbon Dress Size 6US -Very chic! Original Price $525 China Airlines x Moschino Business Class Deluxe Travel Amenity Kit French Blue Moschino Cheap And Chick NWT Dress IT 46 Abstract Print Dolman Sleeve Belted BOUTIQUE MOSCHINO Dress Women Sz 6 RARE Designer Branded Signature Novelty Print Moschino Couture Logo Budweiser printed cotton-jersey joggers pants SZ 10 read Moschino MOS508 C9A 53 Women Eyeglasses Moschino Jeans Women Medium IT 42 Jacket Floral Black Button Long Sleeve Moschino Boutique black jacket coat Sz 6 MOSCHINO FOREVER SAILING FOR MEN EDT SPLASH 20×4.5 ML. MINIATURE (PACK OF 20) MOSCHINO Sunglasses (original Retail Price Was $270.00) MOSCHINO BLACK GRAPHIC PRINT LEATHER CLUTCH RETAIL MADE IN ITALY BNWT MOSCHINO Milano Second Bag Pouch Denim Blue Authentic Used Very Good Item Japan Moschino “I Love 70” Watch New Toy 2 Bubble Gum by 3.4 oz EDT for Women Perfume in Box Moschino This Is Not A Moschino Toy Retro Vintage T-Shirt, S-5XL Moschino Pink Signature Tee T-shirt Rare NEW Moschino Vintage Pebbled Leather Lock Travel Handbag Ivory color Rare 90’s Moschino Dress Apricot Sleeveless Fit Flare Bow Hips Sweetheart Neck Cotton 8 Moschino Lettering Logo Calfskin Women’s Belt in Black with Gold Hardware IT 46 Moschino Women’s 189350 One-Piece Patchwork Animal Print Swimsuit Size 1 Love Moschino Dress Womens 4 Black Sweater Dress Long Sleeves Casual Vintage 80’s Franco Moschino Cheap&Chic RARE Happy Things Shirt 16.5 Men NWOT Moschino Couture by Moschino for Women 1.7 oz Deodorant Parfume Natural Spray L’eau Cheap & Chic by Moschino Eau De Toilette Spray 1.7 oz — Love Moschino Black Yellow Dress Club wear Women’s Dress Size US 2 EU 38 NEW Grey Love Moschino Vest Tank Top With Embroidered Print Size GB 10 Moschino Cheap & Chic Wool Beaded Butterfly Collared Sweater Black Sz 8 vintage cheap and chic Moschino Dress Size 6. MOSCHINO x Disney Top Womens Extra Small Short Sleeve Yellow Tee H&M MOSCHINO Moschino 8-Bit ‘This is not a Moschino Toy’ Bear Logo Tee NEW MOSCHINO MOS128/S MVUIR BLUE AUTHENTIC SUNGLASSES W/CASE 57-14 Love Moschino New Solid Red Sheath Sleeveless Fitted Dress 8 NWOT H&MOSCHINO MOSCHINO H&M JACKET PARKA COAT GOLD SIZE XS Moschino baby girl pink cotton teddy bear logo sweatshirt size 12-18 months, 3 Vintage Moschino Cheap Chic Green Large Bow Shift Mini Dress Sz 8 NEW LOVE MOSCHINO BLACK COTTON LOVE & PEACE EMBROIDERY BLOUSE TOP SHIRT,UK 14 MOSCHINO Jeans Vintage Italy Black/White Check Wool Oversized Shirt Size S Vintage 90s y2k Moschino Mare Dress Slinky Embellished US Size 34 Medium MOSCHINO COUTURE Moschino Couture T-shirt Black ZPA0701 Love Moschino Brown Short Sleeve Shirt size L Cheap and Chic Classic by Moschino Eau De Toilette Spray 1.7oz Love Moschino Men’s Airline Logo Short Sleeve T-Shirt NWT Y2K 2000s Love Moschino Pink Silk Lady Bug Mini Dress drop waist ruffles US 8 LOVE MOSCHINO BASEBALL BLACK PANTS STYLE sz IT 42 US 6 NEW RETAIL $354 MOSCHINO CHEAP AND CHIC BOUCLE DRESS 6 CORAL PINK GOLD BUTTONS BOW FRAN FINE Love Moschino Green Ruffle Draped Long Sleeve Shift Dress Size 10 Love Moschino Womens Sequined Tee Shirt Team Moschino Size 6 NWT MOSCHINO WOMEN’S T SHIRT Yellow Flicker Pink Camouflage Logo (U-002) Moschino Cheap & Chic Vintage Mesh Lavender Embroidered Cami Top | IT 44 | UK 12 MOSCHINO COUTURE BLUE JACKET Trompe L’oeil Blazer USA 4 IT38 $2.4K 2021 RUNWAY Vtg Moschino Jeans Mini Dress Cotton Halter Floral Bow Side Zip Black Small Moschino Olive Oyl Silk Scarf Orchard Heart Cottage Designer Italy Vtg MOSCHINO CHEAP & CHIC 90s DA VINCI OLIVE OYL PRINT BLACK DRESS 42 M Moschino Couture Silk Multicolor Watercolor Abstract Floral Peplum Blazer 42 M Moschino Vintage Faux Leather Jacket PVC Blazer Womens US 8 / It 42 MOSCHINO LOVE DRESS SIZE 10 U.K. RARE Love Moschino Love Logo T-Shirt Black Retro Rainbow HTF Size 8 EUC NWT*MOSCHINO Jeans* shirt-blouse*Size 40 (IT)*RED WINE color*Crystals*VINTAGE Moschino Cheap and Chic Sexy White Draped T Shirt 44/10 Moschino H&M DISNEY Daisy Donald Duck Jumper Sweater Dress S MOSCHINO Women’s Beige Tulip Skirt Asymmetric Pencil Skirt Size 8 / 44 / L MOSCHINO Black Calendar Silk Tie ITALY 56”/ 3.5” EC MOSCHINO mens Shirt Vintage 90s All Over Print Made In Italy 2018 MOSCHINO x H&M Hot Pink Disney Minnie Mouse Merino Wool Sweater Dress – S NWT MOSCHINO HOODIE SWEATSHIRT Teddy bears chains cotton white luxury Italy L Boutique Moschino Button Down Sweater Pink 38 BOUTIQUE MOSCHINO PUSSY BOW BLOUSE TOP PINK SIZE 42 NEW WITH TAGS I LOVE LOVE BY MOSCHINO EDT SPRAY 1.0 OZ (WOMEN) NEW IN BOX Boutique Moschino 100% Silk Dress U.K 12 bnwt RARE LOVE MOSCHINO COUTURE CREW NECK SHORT SLEEVE COTTON WHITE DRESS SIZE 4 nwt Moschino MOS130/S 807 BLACK 55/20/140 WOMAN Sunglasses RRP 365$ BNWT Moschino Mare Blue Shirt 52 / L MOSCHINO HEART SHIRT BLOUSE WOMEN’S size 14 Moschino Cheap and Chic Size 8 Strapless Dress Black Cotton Chiffon Overlay Hot! Moschino Toy 2 3pc Set Women Parfum Spy 1.7 oz + Shower Gel + Body Lotion 50% SALE DESIGNER PERFUMES 10 /$19.95 DG M JACOBS MOSCHINO CARTIER+BAG Moschino Jeans Black Tiered Skirt Womens 8 Skirt Belted Short Zipper Moschino Couture Jeremy Scott all over teddy bear policeman pink knit dress US10 Moschino Green Open Toe Floral Detail Shoes Women’s Size 36 1/2 US 5.5 New With Tags Moschino Arm Tape Logo Light Blue Sweatshirt US M (Retail 185$) Mens Pullover Fleece Hoodie